16-001 16-001t.jpg 16-001v.jpg 62 16-001h.jpg 198 Equisetum arvense has classically dimorphic aerial shoots. Reproductive shoots have strobili just reaching maturity (anthesis) as vegetative axes are becoming conspicuous. Photo taken in May in Bayside, Wisconsin Sphenophyta, Equisetales, sporophyte
16-002 16-002t.jpg 16-002v.jpg 46 16-002h.jpg 152 Equisetum arvense reproductive and vegetative shoots. Sporangiophores are readily visible in the strobilus, as is the basal annulus. Photo taken in May in Bayside, Wisconsin Sphenophyta, Equisetales, sporophyte
16-003 16-003t.jpg 16-003v.jpg 146 16-003h.jpg 252 Embryogenesis in Isoetes. Exoscopic embryo gives rise to shoot, leaf, root and foot. The shoot soon emerges through the wall of the gametophyte to establish the young sporophyte. Diagram from Wardlaw, pg. 119 (1955). Lycophyta, Isoetales, embryogenesis
16-004 16-004t.jpg 16-004v.jpg 91 16-004h.jpg 311 Isoetes megagametophyte soon after germination. Swelling of gametophytic tissue occurs along the trilete mark. At this location and along these grooves, archegonia form that will be fertilized by motile sperm, if present. Lycophyta, Isoetales, gametogenesis
16-005 16-005t.jpg 16-005v.jpg 181 16-005h.jpg 339 Lycophyta, Isoetales, megasporangium
16-006 16-006t.jpg 16-006v.jpg 131 16-006h.jpg 493 Lycophyta, Isoetales, megasporangium, microsporangium
16-007 16-007t.jpg 16-007v.jpg 232 16-007h.jpg 391 Lycophyta, Isoetales, megasporangium
16-008 16-008t.jpg 16-008v.jpg 284 16-008h.jpg 548 Lycophyta, Isoetales, megasporangium
16-009 16-009t.jpg 16-009v.jpg 330 16-009h.jpg 577 Lycophyta, Isoetales, microsporangium
16-010 16-010t.jpg 16-010v.jpg 316 16-010h.jpg 568 Lycophyta, Isoetales, microsporangium
16-011 16-011t.jpg 16-011v.jpg 349 16-011h.jpg 648 Lycophyta, Isoetales, shoot axis
16-012 16-012t.jpg 16-012v.jpg 324 16-012h.jpg 695 Lycophyta, Isoetales, shoot axis
16-013 16-013t.jpg 16-013v.jpg 169 16-013h.jpg 726 Lycophyta, Isoetales, shoot axis
16-014 16-014t.jpg 16-014v.jpg 345 16-014h.jpg 671 Lycophyta, Isoetales, shoot axis
16-015 16-015t.jpg 16-015v.jpg 127 16-015h.jpg 280 Lycophyta, Isoetales, leaf
16-016 16-016t.jpg 16-016v.jpg 108 16-016h.jpg 389 Lycophyta, Isoetales, Stigmarian root
16-017 16-017t.jpg 16-017v.jpg 102 16-017h.jpg 316 Lycophyta, Isoetales, Stigmarian root
16-018 16-018t.jpg 16-018v.jpg 288 16-018h.jpg 541 Lycophyta, Isoetales, shoot axis
16-019 16-019t.jpg 16-019v.jpg 413 16-019h.jpg 842 Lycophyta, Isoetales, rhizotaxis
16-020 16-020t.jpg 16-020v.jpg 162 16-020h.jpg 663 Lycophyta, Isoetales
16-021 16-021t.jpg 16-021v.jpg 188 16-021h.jpg 644 Lycophyta, Isoetales
16-022 16-022t.jpg 16-022v.jpg 185 16-022h.jpg 589 Lycophyta, Isoetales
16-023 16-023t.jpg 16-023v.jpg 198 16-023h.jpg 706 Lycophyta, Isoetales
16-024 16-024t.jpg 16-024v.jpg 372 16-024h.jpg 695 Lycophyta, Isoetales
16-025 16-025t.jpg 16-025v.jpg 382 16-025h.jpg 666 Lycophyta, Isoetales, shoot axis
16-026 16-026t.jpg 16-026v.jpg 174 16-026h.jpg 768 Lycophyta, Isoetales, shoot axis, root apex
16-027 16-027t.jpg 16-027v.jpg 441 16-027h.jpg 899 Lycophyta, Isoetales, shoot axis
16-028 16-028t.jpg 16-028v.jpg 328 16-028h.jpg 616 Lycophyta, Isoetales, plant habit
16-029 16-029t.jpg 16-029v.jpg 157 16-029h.jpg 535 Lycophyta, Isoetales, plant habit
16-030 16-030t.jpg 16-030v.jpg 94 16-030h.jpg 317 Lycophyta, Isoetales, plant habit
16-031 16-031t.jpg 16-031v.jpg 198 16-031h.jpg 344 Lycophyta, Isoetales, plant habit
16-032 16-032t.jpg 16-032v.jpg 146 16-032h.jpg 461 Lycophyta, Selaginellales, megasporangium
16-033 16-033t.jpg 16-033v.jpg 250 16-033h.jpg 440 Lycophyta, Selaginellales, megasporangium
16-034 16-034t.jpg 16-034v.jpg 60 16-034h.jpg 246 Lycophyta, Selaginellales, megasporangium
16-035 16-035t.jpg 16-035v.jpg 185 16-035h.jpg 329 Lycophyta, Selaginellales, megasporangium
16-036 16-036t.jpg 16-036v.jpg 170 16-036h.jpg 286 Lycophyta, Selaginellales, megasporangium
16-037 16-037t.jpg 16-037v.jpg 209 16-037h.jpg 356 Lycophyta, Selaginellales, megasporangium
16-038 16-038t.jpg 16-038v.jpg 96 16-038h.jpg 349 Lycophyta, Selaginellales, microsporangium
16-039 16-039t.jpg 16-039v.jpg 137 16-039h.jpg 399 Lycophyta, Selaginellales, microsporangium
16-040 16-040t.jpg 16-040v.jpg 128 16-040h.jpg 2339 Lycophyta, Selaginellales, microsporangium
16-041 16-041t.jpg 16-041v.jpg 243 16-041h.jpg 1726 Lycophyta, Selaginellales, shoot apex, dichotomous branching
16-042 16-042t.jpg 16-042v.jpg 125 16-042h.jpg 2025 Lycopodium embryogenesis shows the characteristic young leaves of this plant. The classical embryo is endoscopic, with the suspensor located under the archegonial neck and the shoot emerging laterally. Lycophyta, Lycopodiales, embryogenesis
16-043 16-043t.jpg 16-043v.jpg 209 16-043h.jpg 1658 Lycopodium embryogenesis shows a high diversity of different embryogenetic pathways, which is presumed to be a primitive characteristic. The classical embryo is endoscopic, with the suspensor located under the archegonial neck and the shoot emerging laterally. Lycophyta, Lycopodiales, embryogenesis
16-044 16-044t.jpg 16-044v.jpg 128 16-044h.jpg 2030 Lycopodium gametophyte with archegonia show the onset of the sexual phase of this long-lived plant. Note the axial row present in the archegonial neck. Lycophyta, Lycopodiales, gametophyte, archegonium, embryogenesis
16-045 16-045t.jpg 16-045v.jpg 120 16-045h.jpg 2027 Lycopodium gametophyte with mature archegonia showing the axial row in the archegonial neck, ventral canal cell and interior egg cell. Lycophyta, Lycopodiales, gametophyte, archegonium
16-046 16-046t.jpg 16-046v.jpg 130 16-046h.jpg 1975 Lycopodium gametophyte with endophytic fungus, which is characteristic of the slowest growing of the Lycopod gametophytes. The gametophytes of some species may require up to seven years to mature. Lycophyta, Lycopodiales, gametophyte
16-047 16-047t.jpg 16-047v.jpg 103 16-047h.jpg 1912 Lycopodium gametophyte with mature antheridia. The antheridia are embedded in this species, with a single layer of cells at the aperture. Antheridia are typically on the underside of the gametophyte. Lycophyta, Lycopodiales, gametophyte, antheridia
16-048 16-048t.jpg 16-048v.jpg 299 16-048h.jpg 1878 Phylloglossum drummondii stobilus. This image of this unusual plant shows a longitudinal section of the cone, displaying a wide range of stages of sporogenesis. Lycophyta, Lycopodiales, sporangia
16-049 16-049t.jpg 16-049v.jpg 129 16-049h.jpg 1990 Lycopodium obscurum mature eusporangium. Spores continue to increase in density as they accumulate food stores. The tapetum breaks down as its nutrients dry up in preparation for dehiscence. Note the laminar location of sporangial attachment on the sporophyll. Lycophyta, Lycopodiales, sporangium
16-050 16-050t.jpg 16-050v.jpg 258 16-050h.jpg 1938 Lycopodium lucidulum mature eusporangium. Note the axillary location of sporangial attachment. This is characteristic of some of the less derived of the Lycopodiaceae. Lycophyta, Lycopodiales, sporangium
16-051 16-051t.jpg 16-051v.jpg 116 16-051h.jpg 1534 Lycopodium mature eusporangium. Spores continue to increase in density as they accumulate food stores. The tapetum breaks down as its nutrients dry up in preparation for dehiscence. Note the laminar location of sporangial attachment on the sporophyll. This is characteristic of some of the more derived of the Lycopodiaceae. Lycophyta, Lycopodiales, sporangium
16-052 16-052t.jpg 16-052v.jpg 116 16-052h.jpg 1968 Lycopodium clavatum sporogenesis IV. Spores with conspicuous exines separate. Occasionally cells with their trilete mark evident are visible. Tapetal cells found among the sporocytes continue to break down. Lycophyta, Lycopodiales, sporogenesis
16-053 16-053t.jpg 16-053v.jpg 102 16-053h.jpg 1587 Lycopodium clavatum sporogenesis III. Tetrads have broken down, releasting the spores as individual cells. Exine walls are well developed in the spores at this stage. Tapetal cells found among the sporocytes are beginning to break down. Lycophyta, Lycopodiales, sporogenesis
16-054 16-054t.jpg 16-054v.jpg 84 16-054h.jpg 1506 Lycopodium clavatum sporogenesis II. Post-meiotic spores remain in tetrads in this image. Tapetal cells are found among the sporocytes at this stage. Lycophyta, Lycopodiales, sporogenesis
16-055 16-055t.jpg 16-055v.jpg 102 16-055h.jpg 1667 Lycopodium clavatum sporogenesis I. Sporocytes have just undergone meiosis and spores are present in tetrads in this image. Note the thickness of the sporangial wall and the organization of tapetal cells. Lycophyta, Lycopodiales, sporogenesis
16-056 16-056t.jpg 16-056v.jpg 177 16-056h.jpg 1144 Lycopodium late sporangial development. Sporogenous tissue is quite well defined at this stage. The thick walls surrounding developing sporocytes is characteristic. Lycophyta, Lycopodiales, sporogenesis
16-057 16-057t.jpg 16-057v.jpg 213 16-057h.jpg 1664 Lycopodium later sporangial development. Sporogenous tissue is just beginning to become conspicuous by its greater cytoplasmic density and accumulation of dense nuclei. Lycophyta, Lycopodiales, sporogenesis
16-058 16-058t.jpg 16-058v.jpg 111 16-058h.jpg 1830 Lycopodium sporangial development. This eusporangium is still reasonably early in development. Note the origin of layering in the sporangium -- an early stage in formation of the sporangial wall. Lycophyta, Lycopodiales, sporangial ontogeny
16-059 16-059t.jpg 16-059v.jpg 109 16-059h.jpg 2109 Lycopodium sporangial initials are characteristic of the eusporangiate development. Note the axial origin of the sporangium. Lycophyta, Lycopodiales, sporangial ontogeny
16-060 16-060t.jpg 16-060v.jpg 193 16-060h.jpg 347 Phylloglossum drummondii tuber and shoot apex. This diminutive close relative of the genus Lycopodium, believed to be a derived member of the Lycopodiales, is quite rare. Slide made at University of Oklahoma Lycophyta, Lycopodiales, tuber
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